This page is devoted to list all publications related to ENFA or Biomapper. Should you find any reference that should appear here, please let me know.

  For easier reference, I have classified the publications into several categories:

The references are sorted by date, the most recent at the top of each list. 

You may also want to look at my list of publications

ENFA-related theory


  1. Braunisch V., Bollmann K., Graf R. F., Hirzel A.H. (2008). Living on the edge -- Modelling habitat suitability for species at the edge of their fundamental niche. Ecological Modelling 214, 153-167. PDF (1.12 Mb)
  2. Hirzel, A. H. & Le Lay G. (2008). Habitat suitability modelling and niche theory. Journal of Applied Ecology, 45, 1372-1381. PDF

  3. Hirzel, A. H. (2008). Using relative capacity to measure habitat suitability. Israel Journal of Ecology and Evolution, 54, 421-434. PDF

  4. Calenge, C., Darmon, G., Basille, M., Loison, A. & Jullien, J.M. (2008) The factorial decomposition of the Mahalanobis distances in habitat selection studies. Ecology, 89(2), 555-66. PDF (1.49 Mb)

  5. Chefaoui, R.M., Lobo, J.M. 2008. Assessing the effects of pseudo-absences on predictive distribution model performance. Ecological Modelling 210:478-486. PDF (547 Kb)

  6. Hirzel, A. H., G. Le Lay, V. Helfer, C. Randin, and A. Guisan. 2006. Evaluating the ability of habitat suitability models to predict species presences. Ecological Modelling 199:142-152. PDF (969 Kb)

  7. Hirzel, A.H. & Arlettaz, R. (2003) Modelling habitat suitability for complex species distributions by the environmental-distance geometric mean. Environmental Management, 32, 614-623. PDF (1.09 Mb)

  8. Hirzel A.H., Hausser J., Chessel D. & Perrin N. (2002) Ecological-niche factor analysis: How to compute habitat- suitability maps without absence data? Ecology, 83, 2027-2036. PDF (1.01 Mb)

  9. Hirzel, A.H., Helfer, V., Métral, F., 2001. Assessing habitat-suitability models with a virtual species. Ecological Modelling. 145, 111-121. PDF (322 Kb)

Applications of ENFA

International journals


  1. Jiménez-Valverde, A., Gómez, J.F., Lobo, J.M., Baselga, A. & Hortal, J. (2008) Challenging species distribution models: the case of Maculinea nausithous in the Iberian Peninsula. Annales Zoologici Fennici, 45, 200-10. PDF (619 Kb)
  2. Rivera, J.H.V., Huerta, M.A.O. & Guerrer, R. (2008) Analysis of the distribution of Orange-breasted Bunting (Passerina leclanchetii): An endemic species of Mexico's Pacific slope. Ornitologia Neotropical, 19(2), 265-74. PDF (247 Kb, in Spanish)
  3. Mertzanis G, Kallimanis AS, Kanellopoulos N, et al. (2008) Brown bear (Ursus arctos L.) habitat use patterns in two regions of northern Pindos, Greece - management implications. Journal of Natural History 42, 301-315.
  4. Strubbe, D. & Matthysen, E. (2008) Predicting the potential distribution of invasive ring-necked parakeets Psittacula krameri in northern Belgium using an ecological niche modelling approach. Biol Invasions, on-line. PDF (341 Kb)
  5. Allouche, O., Steinitz, O., Rotem, D., Rosenfeld, A. & Kadmon, R. (2008) Incorporating distance constraints into species distribution models. Journal of Applied Ecology, 45(2), 599-609. PDF (623 Kb)
  6. Long, P.R., Zefania, S., Ffrench-Constant, R.H. & Szekely, T. (2008) Estimating the population size of an endangered shorebird, the Madagascar plover, using a habitat suitability model. Animal Conservation, 11(2), 118-27. PDF (466 Kb)
  7. Praca, E. & Gannier, A. (2008) Ecological niches of three teuthophageous odontocetes in the northwestern Mediterranean Sea. Ocean Science, 4(1), 49-59. PDF (1.73 Mb)
  8. Skov, H., Humphreys, E., Garthe, S., Geitner, K., Gremillet, D., Hamer, K.C., Hennicke, J., Parner, H. & Wanless, S. (2008) Application of habitat suitability modelling to tracking data of marine animals as a means of analyzing their feeding habitats. Ecological Modelling, 212(3-4), 504-12. PDF (987 Kb)
  9. 2007

  10. Chefaoui, R.M., Lobo, J.M. 2007. Assessing the conservation status of an iberian moth using pseudo-absences. Journal of Wildlife Management 71:2507-2516. PDF (152 Kb)
  11. Estrada-Peña, A., Venzal, J.M. 2007. Climate niches of tick species in the mediterranean region: Modeling of occurrence data, distributional constraints, and impact of climate change. Journal of Medical Entomology 44:1130-1138. PDF (805 Kb)
  12. Sattler, T., Bontadina, F., Hirzel, A.H. & Arlettaz, R. (2007) Ecological niche modelling of two cryptic bat species calls for a reassessment of their conservation status. Journal of Applied Ecology, 44(6), 1188-99. PDF (334 Kb)
  13. Jimenez-Valverde A, Ortuno VM, Lobo JM (2007) Exploring the distribution of Sterocorax Ortuno, 1990 (Coleoptera, Carabidae) species in the Iberian peninsula. Journal of Biogeography 34, 1426-1438. PDF (679 Kb)
  14. Braunisch, V. and Suchant, R. 2007. A model for evaluating the 'habitat potential' of a landscape for capercaillie Tetrao urogallus: a tool for conservation planning. - Wildlife Biology 13: 21-33. PDF (441 Kb)

  15. Soares, C. and Brito, J. C. 2007. Environmental correlates for species richness among amphibians and reptiles in a climate transition area. - Biodiversity and Conservation 16: 1087-1102. PDF (273 Kb)

  16. Titeux, N., Dufrêne, M., Radoux, J., Hirzel, A.H. & Defourny, P. (2007) Fitness-related parameters improve niche-based distribution modelling: the case of the Red-backed Shrike. Biological Conservation, 138, 207-23. (PDF 1.13 Mb)

  17. Mestre, F.M., Ferreira, J.P. & Mira, A. (2007) Modelling the distribution of the European Polecat Mustela putorius in a Mediterranean agricultural landscape. Revue d'Ecologie (Terre Vie), 62, 35-47. (PDF 420 Kb)

  18. Traill, L.W. & Bigalke, R.C. (2007) A presence-only habitat suitability model for large grazing African ungulates and its utility for wildlife management. African Journal of Ecology, 45, 347-354. (PDF 496 Kb)

  19. Wilson, M.F.J., O’Connell, B., Brown, C., Guinan, J.C. & Grehan, A.J. (2007) Multiscale Terrain Analysis of Multibeam Bathymetry Data for Habitat Mapping on the Continental Slope. Marine Geodesy, 30:3-35. (PDF 1.42 Mb)

  20. Bryan, T.L. and Metaxas, A. 2007. Predicting suitable habitat for deep-water gorgonian corals on the Atlantic and Pacific Continental Margins of North America. Marine Ecology Progress Series 330:113-126. PDF (1.23 Mb) 

  21. 2006
  22. Santos, X., Brito, J.C., Sillero, N., Pleguezuelos, J.M., Llorente, G.A., Fahd, S. & Parellada, X. (2006) Inferring habitat-suitability areas with ecological modelling techniques and GIS: A contribution to assess the conservation status of Vipera latastei. Biological Conservation, 130(3), 416-25. PDF (343 Kb)
  23. Cassinello, J., P. Acevedo, and J. Hortal. 2006. Prospects for population expansion of the exotic aoudad (Ammotragus lervia; Bovidae) in the Iberian Peninsula: clues from habitat suitability modelling. Diversity and Distributions 12:666-678. PDF (376 Kb)

  24. Estrad-Peña, A., Venzal, J.M., Acedo, C.S., 2006. The tick Ixodes ricinus: distribution and climate preferences in the western Palaearctic. Medical and Veterinary Entomology 20, 189-197. PDF (775 Kb)

  25. Tole, L., 2006. Choosing reserve sites probabilistically: A Colombian Amazon case study. Ecological Modelling 194, 344-356. PDF (577 Kb)

  26. 2005
  27. Boisteau B., Marion L., 2005. Definition of potential habitats of Grey Heron using landscape analysis and the species ecological niche [Définition des habitats potentiels du héron cendré Ardea cinerea par l'analyse du paysage et de sa niche écologique]. Alauda 73, 431-440. PDF (5.36 Mb)

  28. Chefaoui, R., Hortal, J. and Lobo, J.M. (2005) Potential distribution modelling, niche characterization and conservation status assessment using GIS tools: a case study of Iberian Copris species. Biological Conservation, 122, 327-338. PDF (519 Kb)

  29. 2004
  30. Gallego D, Canovas F, Esteve MA, Galian J, Schwerdtfeger F. 2004. Descriptive biogeography of Tomicus (Coleoptera : Scolytidae) species in Spain. Journal of Biogeography 31(12):2011-2024. PDF (417 Kb)

  31. Hirzel AH, Posse B, Oggier P-A, Crettenand Y, Glenz C, Arlettaz R. 2004. Ecological requirements of a reintroduced species, with implications for release policy: the Bearded vulture recolonizing the Alps. Journal of Applied Ecology 41:1103-1116. PDF (233 Kb)

  32. Brotons, L., Thuiller, W., Araujo, M.B., & Hirzel, A.H. (2004) Presence-absence versus presence-only modelling methods for predicting bird habitat suitability. Ecography, 27, 437-448. PDF (1.05 Mb)

  33. Engler, R., Guisan, A., & Rechsteiner, L. (2004) An improved approach for predicting the distribution of rare and endangered species from occurrence and pseudo-absence data. J Appl Ecology, 41, 263-274. PDF (311 Kb)

  34. 2003
  35. Dettki, H., Löfstrand, R. & Edenius, L. 2003. Modeling habitat suitability for moose in coastal northern Sweden: empirical vs. process-oriented approaches. AMBIO, 32 (8): 549-556. PDF (464 Kb)

  36. Thomas, C. J. 2003. Anopheles gambiae and climate in Brazil. Lancet Infectious Diseases 3:326-326.

  37. Reutter, B.A., V. Helfer, A.H. Hirzel, and P. Vogel. 2003. Modelling habitat-suitability on the base of museum collections: an example with three sympatric Apodemus species from the Alps. Journal of Biogeography 30, 581-590. PDF (263 Kb)

  38. 2002
  39. Zaniewski A.E., Lehmann A, & Overton J. McC. (2002): "Predicting species spatial distributions using presence-only data: a case study of native New Zealand ferns.". Ecological Modelling, 157, 261-280. PDF (571 Kb)

PhD theses

  1. Wilson, Margaret F.J. (2006) Deep sea habitat mapping using a remotely operated vehicle: Mapping and modelling seabed terrain and benthic habitats at multiple scales in the Porcupine Seabight, SW Ireland.  PhD Thesis, National University of Ireland, Galway. 266pp. PDF (30 Mb: Contact Margaret)

  2. Titeux, N., 2006. Modelling species distribution when habitat occupancy departs from suitability: Application to birds in a landscape context. PhD thesis. Université Catholique de Louvain, Belgique. 256pp. PDF (1.95 Mb)

  3. Freer RA. 2004.The spatial ecology of the Güiña (Oncifelis guigna) in Southern Chile. Durham: University of Durham. (PhD thesis) 219 p. PDF (9.46 Mb) 

  4. Zimmermann F. 2004. Conservation of the Eurasian Lynx (Lynx lynx) in a fragmented landscape - habitat models, dispersal and potential distribution [PhD thesis]. Lausanne: University of Lausanne. 180 p. PDF (5.66 Mb)

  5. Williams, A.K. (2003) The Influence of Probability of Detection When Modeling Species Occurrence Using GIS and Survey Data. PhD thesis, Blacksburg University, Blacksburg (USA). PDF(1.62Mb)

  6. Patthey, P. 2003. Habitat and corridor selection of an expanding red deer (Cervus elaphus) population. PhD thesis. Institute of Ecology, University of Lausanne. PDF (4.54 Mb)

  7. Sachot, S., 2002. Viability and management of an endangered capercaillie (Tetrao urogallus) metapopulation. PhD thesis. Institute of Ecology, University of Lausanne. PDF (1.66 Mb).

  8. Hirzel, A.H., 2001. When GIS come to life. Linking landscape- and population ecology for large population management modelling: the case of Ibex (Capra ibex) in Switzerland. PhD thesis. Institute of Ecology, Laboratory for Conservation Biology. University of Lausanne. PDF (1.61 Mb) or cyberdocument.

  9. Perrin, N., 1984. Contribution à l'écologie du genre Cepaea (Gastropoda): Approche descriptive et expérimentale de l'habitat et de la niche écologique. PhD thesis. Institut de Zoologie et d'Ecologie Animale. University of Lausanne.


  1. Olson D.E and Davis R.J. Conservation Assessment for the Foothill Yellow-legged Frog (Rana boylii) in Oregon, Appendix 1. 2007. USDA Forest Service Region 6 and USDI Bureau of Land Management. PDF (Appendix I, 983 Kb) Whole report
  2. Clark MR, Tittensor D, Rogers AD, Brewin P, Schlacher T, Rowden A, Stocks K, Consalvey M (2006). Seamounts, deep-sea corals and fisheries: vulnerability of deep-sea corals to fishing on seamounts beyond areas of national jurisdiction. UNEPWCMC,Cambridge, UK. PDF (1.84 Mb)
  3. Raphael, M. G., B. Galleher, M. H. Huff, S. L. Miller, S. K. Nelson, and R. D. Young. 2006. Spatially-explicit estimates of potential nesting habitat for the Marbled Murrelet. Pages 97-146 in M. H. Huff, R. M.G., M. S.L., N. S.K., and Baldwin, editors. Northwest Forest Plan—the first 10 years (1994-2003): status and trends of populations and nesting habitat for the marbled murrelet. PNW-GTR-650, U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station, Portland, USA. PDF (3.88 Mb)

  4. Lint J., Martin J., Davis R., Mowdy J., Reid J., Forsman E., Anthony R., 2005. Northwest forest plan - The first 10 years (1994-2003): Status and trends of northern spotted owl populations and habitat. USDA Forest Service - General Technical Report PNW, 1-176. PDF (8.22Mb) URL

  5. Breitenmoser, U., Zimmermann, F., Olsson, P., Ryser, A., Angst, C., Jobin, A., Breitenmoser-Würsten, C., 1999. Beurteilung des Kantons St.Gallen als Habitat für den Luchs. Bern, KORA.

Books and book chapters

  1. Hortal J., Borges P.A.V., Dinis F., Jiménez-Valverde A., Chefaoui R.M., Lobo J.M., Jarroca S., Azevedo E.B.d., Rodrigues C., Madruga J., Pinheiro J., Gabriel R., Rodrigues F.C., Pereira A.R., 2005. Using ATLANTIS - TIERRA 2.0 and GIS environmental information to predict the spatial distribution and habitat suitability of endemic species. In: A list of the terrestrial fauna (Mollusca and Arthropoda) and flora (Bryophyta, Pteridophyta and Spermatophyta) from the Azores (Borges PAV, Cunha R, Gabriel R, Martins AF, Silva L, Vieira V(Borges PAV, Cunha R, Gabriel R, Martins AF, Silva L, Vieira Vs. Borges PAV, Cunha R, Gabriel R, Martins AF, Silva L, Vieira V), pp. 69-113. Direcção Regional de Ambiente and Universidade dos Açores, Horta, Angra do Heroísmo and Ponta Delgada. PDF (7.95 Mb) (English and Portuguese)

  2. Hirzel, A.H. & Arlettaz, R. (2003) Environmental-envelope based habitat-suitability models. In 1st Conference on Resource Selection by Animals (ed B.F.J. Manly). Omnipress, Laramie, USA. PDF (2.56 Mb)

  3. Chatelain C., Gautier L. & Spichiger R. (2002) Application du SIG IVOIRE à la distribution potentielle des espèces en fonction des facteurs écologiques. In: Plant systematics and phytogeography for understanding of african biodiversity. Systematics and Geography of Plants 71(2). (ed. Robbrecht E, J. Degreef & I. Friis), Bruxelles. PDF (845Kb).

  4. Hausser, J., 1995. Mammifères de Suisse. Bâles, Birkhäuser.

Master theses

  1. Compton, R. C. 2004. Predicting key habitat and potential distribution of Northern Bottlenose Whales (Hyperoodon ampullatus) in the northwest Atlantic ocean. MSc thesis. University of Plymouth, 40 p. PDF (3.15 Mb)

  2. Mandleberg, L. 2004. A comparison of the predictive abilities of four approaches for modelling the distribution of cetaceans. MSc thesis. School of Biological Sciences, University of Aberdeen, UK, 54p. PDF (2.38 Mb)

  3. Leverette, T. L. 2004. Predicting suitable habitat for deep water corals in the Pacific and Atlantic Continental Margins of North America. MSc thesis. Department of Oceanography. - Dalhousie University, 81 p. PDF (2.49 Mb)

  4. Bassi, M. 2003. Habitat-Modelle in der Wildökologie. Masters. Institut für Wildbiologie und Jagdwirtschaft, Universität für Bodenkultur, Wien. PDF (German) (1.25 Mb)
  5. Saudan, M., and A. Hirzel. 2000. Corrélation entre facteurs écogéographiques et capacité de soutien chez le chevreuil (Capreolus capreolus) au Valais. Masters thesis. Laboratory for Conservation Biology, University of Lausanne. PDF (French) (537 Kb).
  6. Helfer, V., F. Métral, and A. Hirzel. 2000. Construction de cartes d'habitat potentiel : Comparaison et évaluation de l'ENFA et du GLM à l'aide d'une espèce virtuelle : le Marsupilami. Certificate. University of Lausanne. PDF (French) (1.44 Mb). Contact V.Helfer and F.Métral
  1. Hirzel A.H., Le Lay G., Helfer V., Randin C. and Guisan A. 2007. Evaluating presence-only habitat-suitability models. European Conference of Ecological Modelling. PDF (3.03 Mb)

  2. Ortega M. & Václavík T. 2007. Modelling potential distribution of Norway Maple (Acer platanoides) in Massachussets, USA. A comparative assessment of Mahalanobis typicality and Ecological Niche Factor Analysis. Presented at Clark University workshop. PDF (1.70 Mb)
  3. White, E.C. 2006. What makes good quality mandrill (Mandrillus sphinx) habitat? Presented at the ISBE conference, Tours. PDF (2.06 Mb)

  4. Witt, J. and S. Menon. 2004. Conservation of the ‘Vulnerable’ Beautiful Nuthatch, Sitta formosa: A Preliminar Analysis of Species Distribution and Habitat Requirements. Society for Conservation Biology Conference. PDF (215 Kb)

  5. Hirzel, A.H., J. Hausser and N. Perrin. 2000. How to compute a Habitat Suitability map without absence data? Use the Ecological Niche Factor Analysis (ENFA)! PDF (138 Kb)
Related papers
  1. Carsten, F. Dormann, Jana, M. McPherson, Miguel, B. Araujo, Roger, Bivand, Janine, Bolliger, Gudrun, Carl, Richard, G. Davies, Alexandre, Hirzel, Walter, Jetz, Kissling, W. Daniel, Ingolf, Kühn, Ralf, Ohlemüller, Pedro, R. Peres-Neto, Björn, Reineking, Boris, Schröder, Frank, M. Schurr, Robert, Wilson, 2007. Methods to account for spatial autocorrelation in the analysis of species distributional data: a review. Ecography 30, 609-628. PDF (716 Kb)

  2. Colwell, R.K. & Futuyma, D.J. (1971) Measurement of Niche Breadth and Overlap. Ecology, 52(4), 567-76. (PDF 1.46 Mb)

  3. Guisan, A., Zimmermann, N.E., 2000. Predictive habitat distribution models in ecology. Ecological Modelling. 135, 147-186. PDF (1.05 Mb)
  4. Hurlbert, S.H. (1978) The measurement of niche overlap and some relatives. Ecology, 59(1), 67-77. (PDF 1.43 Mb)

  5. Hutchinson, G.E. (1957) Concluding remarks. In Cold Spring Harbour symposium on quantitative biology, Vol. 22, pp. 415-427. PDF (1.09 Mb)

  6. Lobo, J.M. 2007. More complex distribution models or more representative data? Biodiversity Informatics 4.
     PDF (413 Kb)
  7. Lobo, J.M., Jimenez-Valverde, A., Real, R. 2007. Auc: A misleading measure of the performance of predictive distribution models. Global Ecology and Biogeography. on-line. PDF (139 Kb)
  8. Peterson, A.T. 2006. Uses and requirements of ecological niche models and related distributional models. Biodiversity Informatics 3:59-72. PDF (114 Kb)
  9. Soberon, J. 2007. Grinnellian and eltonian niches and geographic distributions of species. Ecology Letters 10:1115-1123. PDF (802 Kb)

My other publications


  1. Niculita-Hirzel H., Hirzel A. H., 2008. Visualizing the gene order conservation among genomes with ChromoMapper. International Journal of Computational Intelligence in Bioinformatics and Systems Biology 1(2). in press
  2. Hirzel, A.H., Nisbet, R.M. & Murdoch, W.W. (2007) Host-parasitoid spatial dynamics in heterogeneous landscapes. Oikos, 116:2082-2096. PDF (474 Kb)
  3. Hirzel A.H. (2006) : “Les profondeurs de Chillon”. In “Si une Suisse fantastique m’était contée”, ed. Zoé. Pp 77-91.

  4. Arditi R., Michalski J., Hirzel A.H., 2005. Rheagogies: Modelling non-trophic effects in food webs. Ecological Complexity 2, 249-258. PDF (272 Kb) 

  5. Hirzel, A., and A. Guisan. 2002. Which is the optimal sampling strategy for habitat suitability modelling. Ecological Modelling 157:331-341. PDF (178 Kb)
  6. Hirzel, A., and N. Perrin. 1998. Modeling the dynamics and management of Ibex populations in Switzerland. Pages 139-144 in Proc. 2nd World Conf. Mt. Ungulates. PDF (82 Kb)

  7. Hirzel A. 2000. Terra Australis Incognita. in Revue Archipel 18, 61-86. Revue Archipel

  8. Hirzel A. 1997. La faille. in Revue Archipel 13, 51-69. Revue Archipel
  9. Hirzel A. 1996. Chillon. in Revue Archipel 11, 77-105. Revue Archipel

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